A biologist spent many years researching the rate of evolutionary change in the finch populations of a group of islands. It was determined that the average beak size (both length and mass) of finches in a certain population increased dramatically during an intense drought between 1981 and 1987. During the drought, there was a reduction in the number of plants producing thin-walled seeds. The biologist discovered that from 1988 to 1993, the average beak size declined to pre-1981 levels. The reversal in beak size from 1988 to 1993 was most likely related to which of the following events?